The bands of riparian vegetation (arboreal, shrubby or herbaceous) play such an important role that they become inseparable from the river. When left to their natural dynamics, these environments are characterized by their remarkable dynamism and heterogeneity of habitats, and are able to support a high biodiversity. Periodic floods are exploited by aquatic and terrestrial organisms to increase diversity and productivity; they are, in fact, generally more productive environments then their surrounding territories.
The plants are selected mainly by the water regime that affects soil texture, water availability and nutrient replenishment. On the edge of the shore, sometimes preceded by populations of herbaceous plants and marsh reeds immersed in water, we almost always find thick bushes of shrubby willows (Salix cinerea, Salix purpurea and Salix eleagnos) followed by a row or two of Poplars (Populus nigra) and tree willows (Salix alba). These forest groups, along with the alder wood in the finer soils, typical of the riparian areas, gradually turn into mature woods composed of ash trees, elms, rural maples, oaks, hornbeams etc
The white willow, is a species of willow native to Europe and western and central Asia. The name derives from the white tone to the undersides of the leaves. It is a medium-sized to large deciduous tree growing up to 10–30 m tall, with a trunk up to 1 m diameter and an irregular, often-leaning crown. The bark is grey-brown, and deeply fissured in older trees. The shoots in the typical species are grey-brown to green-brown. The leaves are paler than most other willows, due to a covering of very fine, silky white hairs, in particular on the underside; they are 5–10 cm long and 0.5–1.5 cm wide. The flowers are produced in catkins in early spring, and pollinated by insects. It is deciduous, with male and female catkins on separate trees; the male catkins are 4–5 cm long, the female catkins 3–4 cm long at pollination, lengthening as the fruit matures. When mature in midsummer, the female catkins comprise numerous small (4 mm) capsules, each containing numerous minute seeds embedded in white down, which aids wind dispersal. he white willow mainly grows in temperate climates on spots with abundant light. Its roots need access to water; hence, the willow is usually found in riparian ecosystems. It tolerates a wide variety of soils, but prefers those which are sandy and calcareous. The tree occurs in altitudes up to 2400 m.
The black poplar is a pioneer deciduous wind-pollinated tree species, widely distributed across Europe, Asia and northern Africa. It reaching heights of up to 40m tall and trunk diameters of up to 200cm. The bark is dark brown or black, with numerous fissures. The leaves are variable in size and shape, longer than wider, but they usually have a cuneate base and serrated margins. The flowers appear before the foliage develops, from specialised buds containing preformed inflorescences. The fruits consist of capsules grouped in catkins. It can be propagated both in generative (by wind- and water-dispersed seeds) and vegetative ways (by cuttings). There are a large number of clones, varieties and hybrids, making classification difficult. Mature trees can live for 100, occasionally 300-400 years. It is a tree species of floodplain forests, growing in riparian mixed forests together with white poplar (Populus alba L.), willow (Salix spp.), alder (Alnus spp.), maple (Acer spp.), elm (Ulmus spp.), and sometimes oak (Quercus spp.). It is a pioneer tree species and does not tolerate drought or shade. It is an opportunistic species able to colonise new sites after disturbances, and has a good tolerance to high water levels and high temperatures during summer.
It is a fast-growing and fairly short-lived tree that grows up to 10–25 m (35–80 ft) tall, with a trunk diameter of 30–50 cm (12–20 in)}, rarely up to 1 m (3.3 ft)} diameter. It often has several trunks and can form impenetrable thickets.[4] The typical lifespan of box elder is only 60 years. Under exceptionally favorable conditions, it may live to 100 years. The shoots are green, often with a whitish to pink or violet waxy coating when young. Branches are smooth, somewhat brittle, and tend to retain a fresh green color rather than forming a bark of dead, protective tissue. The bark on its trunks is pale gray or light brown, deeply cleft into broad ridges, and scaly. It has pinnately compound leaves that usually have three to seven leaflets. Simple leaves are also occasionally present; technically, these are single-leaflet compound leaves. The yellow-green flowers are small and appear in early spring, with staminate flowers in clusters on slender pedicels and pistillate flowers on drooping racemes 10–20 cm long. The fruits are paired samaras on drooping racemes, each seed slender, 1-2 cm long ncurved wing; they drop in autumn or they may persist through winter. Seeds are usually both prolific and fertile. Unlike most other maples, A. negundo is fully dioecious and both a male and female tree are needed for reproduction to occur. In Europe where it was introduced in 1688 as a park tree it is able to spread quickly and is considered an pest invasive species in parts of Central Europe, including Germany, the Czech Republic, Hungary, Slovenia, and Poland) where it can form mass growth in lowlands, disturbed areas, and riparian biomes on calcareous soils. This species prefers bright sunlight. It often grows on flood plains and other disturbed areas with ample water supply, such as riparian habitats.Human influence has greatly favored this species; it grows around houses and in hedges, as well as on disturbed ground and vacant lots. It is found on fallow lands and along rivers, where the plant is able to establish in artificial banks. Box-elder is reported to prefer moist, eutrophic and warm conditions. It readily colonises resource-rich riparian forests in Central and Southern Europe (e.g. Salicion albae and Alno-Padion). Young trees prefer moist sites, but become drought tolerant once well established. Seeds are easily dispersed by wind and running water over long distances. In Central Europe, box-elder is tolerant to flooding and is especially competitive in wide floodplains where vegetative regrowth can be impressive. However, in the absence of flooding, it is usually replaced in the course of succession by more shade-tolerant species. In optimal conditions, its permanent and fast vegetative regrowth leads to the formation of dense clones and to the exclusion of other tree species (Salix alba) and herbaceous plants. It favours replacement of softwood by hardwood tree species and makes a threat for this endangered habitat. A more invasive behaviour is however likely to be observed in the future due to climate warming.
Rubus caesius is a Eurasian species of dewberry, known as the European dewberry. Like other dewberries, it is a species of flowering plant in the rose family, related to the blackberry. It is widely distributed across much of Europe and Asia Rubus caesius is similar to and often confused with forms of Rubus fruticosus. It is a small shrub growing up to 2 m tall with biennial stems which die after fruiting in their second year. It sends out long runners which root at the tip to form new plants. The stems are bluish-grey and sometimes prickly. The alternate leaves are hairy above and below. They are stalked and the leaf blades are palmate in shape, either consisting of three oval leaflets with serrated margins and acute points or just being three-lobed. The inflorescence is a loose cluster of several white flowers about 2.5 cm in diameter. The calyx has five sepals and the corolla is composed of five spreading petals with finely toothed margins.
The black alder is a short-lived (around 60 years with a maximum of up to 160 depending on the region), rather small (between 10 and 25m tall) but fast growing broadleaved tree that can be found over most of Europe. It needs a high availability of moisture to grow well, and can often be found along river banks, lake shores and in marshy locations. The roots are well-adapted to growing on very wet soils: it can survive flooding better than most other forest tree species. It is able to fix nitrogen in symbiotic root nodules making it useful for improving soil condition. The timber is durable under water and is often used for jetties and underwater supports, for example in Venice. The bark is brown and smooth at first, becoming darker, rough and fissured with age. The dark green leaves are simple, obovate and measure 4-10cm. Flowering starts before bud burst. The young buds are sticky, giving rise to the name “glutinosa”. Alder is monoecious and the male and female catkins develop in the autumn of the previous year, appearing early in the following spring. The fruits are woody and resemble small pine cones. After wind pollination the seeds, which float well owing to their corky float chambers and oily water-resistant outer coat, are mainly dispersed by water. It is often found together with ash (Fraxinus spp.), birch (Betula spp.), willow (Salix spp.) or oak (Quercus spp.).
The White poplar is a medium-sized tree commonly occurring in coastal and riparian forests of central and southern Europe. It is a fast-growing pioneer tree, which thrives in borders and sunny habitats in sandy alluvial soils and dunes. This poplar covers an important ecological role as a component of floodplain mixed forests, which are ecosystems with very high biodiversity and that are strongly threatened by human activities This poplar dominates and co-dominates in riparian woodland of the central-western Mediterranean zone with willows (Salix alba, Salix fragilis), black poplar (Populus nigra), and alder (Alnus glutinosa). In the wooded steppe zone it is dominant on sandy soils having originated from alluvial deposits and on sand dunes bordering riverine gallery forests . It grows in most sites, tolerating from waterlogged to drought habitats and from acid to strong alkaline soils, but developing in shrub form in extreme conditions. New plants may also establish through tree fragments, which can easily root in suitable environments. Suckers from a single tree can quickly develop a dense and large colony, shading out competitive vegetation. Above or in young trees the bark is creamy white pitted with small black diamonds; it is black and coarsely cracked at the base of older trees. The leaves are alternate, morphologically variable, with 3-5 lobes coarsely toothed, 6-12cm long and longer than they are wide. The colour is shiny dark-green on the upper side and white with dense hair on the lower side. Flowers are out before the leaves in early spring. The male catkins are grey with red stamens, 5-8cm long; the female catkins are greyish-green, 10-15cm long, forming fluffy seeds in early summer. Mature trees also produce abundant wind-dispersed seeds that may be carried long distances. The white poplar hybridises naturally with Eurasian aspen (Populus tremula). The resulting hybrids are known as grey poplars (Populus x canescens), which is a morphologically intermediate species, but which exhibits more vigour than either parent.
Black locust is a tree native to North America and is one of the most important and widespread broadleaved alien trees in Europe. It is a medium-sized, deciduous, fast-growing thorny tree with high suckering capacity. It has been extensively planted in Europe and now it is naturalised in practically the whole continent. It tolerates a remarkably diverse range of soil conditions, being only limited by low soil aeration and waterlogging. It is mainly distributed in sub-Mediterranean to warm continental climates and requires a rather high heat-sum. As a light-demanding pioneer species, it rapidly colonises grasslands, semi-natural woodlands and urban habitats, where it can persist for a long time. Owing to the capacity of fixing di-nitrogen through symbiotic rhizobia in root nodules, black locust can add high rates of nitrogen to soil and change its conditions. The leaves are composed, pinnate, 10-30cm long, usually with a pair of spines at the base which persist on young shoots. The leaflets are commonly in 2-12 pairs, usually opposite, with an additional one at the end of the rachis, white and fragrant flowers are arranged in pendulous racemes 10- 20cm long. The fruit is a legume, 5-10cm long dark brown pods hanging in winter and containing 4-10 seeds, mainly dispersed by gravity and wind .. Climate warming is expected to favour its further expansion. Its reproduction is primary asexual through horizontal root elongation22. It is able to produces a high number of root suckers and therefore exhibits the capacity for clonal growth, up to an area of 100 square metres. Disturbance favours clonal growth and causes an increase in the number of suckers. The invasion of black locust is well documented in early succession habitats. Its adaptability and the capacity of transforming ecosystem processes are the reason for its adverse effects on biodiversity. However, inferior competitive ability in later successional stages is reported where stands have been left unmanaged for a sufficient time.
The tree of heaven is native to China, but it can be found in several countries across Europe and North America. Tree-of-Heaven is known as an invasive species that can rapidly spread onto disturbed sites or fragmented landscapes. It is readily propagated both by seed and vegetatively. It is limited by low temperatures in the north but is abundant across the Mediterranean region. Its expansion has been facilitated by the worldwide transfer of seeds over the last two centuries and by its ability to grow on poor sites, urban areas and fragmented landscapes. It is a short lived, fast-growing tree, reaching a height of around 20m and 70cm in diameter. The bark is greyish and slightly rough. The leaves are 0.4-0.7m long; each leaf comprises 11-25 ovatelanceolate leaflets which are 5-10cm long. The leaflets usually have one or more coarse teeth at the base and each of these teeth has a characteristic gland on the underside. The flowers appear in late spring, the trees being usually dioecious, but in some cases both sexes exist on the same individual. Male plants have a foul odour while flowering. The winged fruits are twisted at the top, membranous, purplish yellow and up to 5cm long. Tree of heaven tolerates a wide variety of soil types and climatic conditions. It demands a warm climate, but is resistant to drought and air pollution although it is sensitive to ozone. It is a shade intolerant species, preferring open spaces. The species is known for its ability to produce allelopathic compounds in its leaflets and bark which are toxic to numerous species and which may have potential for development as a natural herbicide. However, its pollen is a known allergen, and its invasive nature means that it is currently in the top 20 environmental weeds identified as targets of classical biological control in Europe.
Fraxinus angustifolia Vahl, narrowed-leaved ash, is a medium-sized tree growing 40-45m in height and up to 1.5m in diameter. The crown is dense, irregular and dome shaped, with short and pendulous shoots. Its bark is grey and becomes finely and deeply reticulate-fissured. The leafs are compound, arranged in groups of 7-13, odd pinnate, and are slender, 3-8cm long and 1-1.5cm broad, shiny green and hairless. The species is monoecious with hermaphrodite inflorescences of 10-30 flowers. Flowers are wind pollinated, developing in early spring or even in autumn, without petals, green with dark purple stigmas and anthers. The fruit is a samara 3-4cm long, flattened, with a distal wing, ripening at the end of the summer. Its distribution covers the central-southern Europe and northwest Africa, up to the Caucasus The northern part of its distribution overlaps with that of common ash (Fraxinus excelsior), with which it can naturally hybridise, developing plants with intermediate trait forms. It is a fast growing pioneer species, occurring in temperate forests. In Mediterranean regions it tends to grow in cooler areas at higher elevations or along rivers and wetlands. If can be found as dominant or secondary species in mixed broadleaved forests. Similar to common ash, it is susceptible to dieback caused by the fungus Chalara fraxinea, which has caused serious damage in nurseries and in lowland forest stands of central Europe. It is a fast-growing and light-demanding tree, occurring mostly in temperate mild climates, with an annual precipitation between 400 and 800mm. It grows well on moist soils, in temporary flooded lowlands, ash occurs mainly in the lowlands, in riparian and floodplain forests along large rivers, where it formed vast and continuous populations, now with more limited extent. In the Mediterranean regions its distribution is more patchy and reduced to smaller and more isolated populations along rivers or on wetland sites, but also on drier sites at higher altitudes. Rarely, it creates pure stands, found only in optimum conditions as a pioneer species. More often this ash is a dominant or a secondary species in mixed broadleaved forests with oak (Quercus spp.), elm (Ulmus spp.), maple (Acer spp.), willow (Salix spp.), poplar (Populus spp.) and lime (Tilia spp.), forming stable and species-rich ecosystems.
Elms are deciduous medium-sized trees growing up to 20m, characterised by pollination and seed dispersal driven by the wind. They are a component of cool mixed broadleaved forests often associated with noble trees such as ashes (Fraxinus spp.), maples (Acer spp.) and limes (Tilia spp.). They show a clear preference for water- and nutrient-rich soils, distributed principally near rivers and streams or on floodplains. Despite this it can tolerate moderately dry and deep soils, preferring slightly acid and siliceous soils. The hairs on the samaras increase their ability to float by surface tension, favouring long-distance dispersal by water. The leaves vary greatly in dimension and form. They are alternate, dark green, usually asymmetrical at the base and acuminate at the apex, with toothed margins. Elms are monoecious hermaphrodite species: the flowers appear before the leaves in early spring, arranged in clusters of 10-30 elements, close to the shoot in U. glabra and U. Minor. The wind pollinated flower is bellshaped, without petals, and dark purplish-red to brownish. The fruits are ovate samaras, with a single central nut seed rounded by a membrane. These tree species have always played an important cultural role, being part of the traditional rural landscape as a tree of multiple purposes, such as working and firewood, fodder supplier, living grapevine support and more recently as an ornamental and roadside tree. The introduction into Europe in the last century of the fungal pathogen Ophiostoma, causing the Dutch elm disease, has devastated the elm populations throughout Europe with two destructive epidemics. Despite their drastic reduction, elms are not considered endangered species, as the main losses regarded mature cultivated trees in urban areas and countryside. Breeding programmes have selected hybrids resistant to Dutch elm disease for ornamental purposes, while in natural forests elms are less susceptible to this pathogen and more threatened by habitat reduction and water regulation.
The False indigo is a shrub native to North America which has been cultivated mainly for its ornamental features, honey plant value and protective properties against soil erosion. It is registered amongst the most noxious invasive species in Europe. It is typically found growing in moist open woodlands, roadsides, canyons, floodplains, gravel bars, stream and pond banks, and along swamp edges. The genus name Amorpha is Greek for amorphos which means formless or deformed, and refers to the flowers which having only one petal are very different from other members of the pea family. Plant in the full sun to partial shade moist to dry soil. It is tolerant of a large range of soil types. The pinnately compound leaves are clustered on the upper third of the plant and give it a fine texture and an airy appearance. It spreads easily by seeds and suckers, and can form dense thickets. The horizontal growth creates a situation where its spread will often be greater than its height. The False indigo is a shrub with a pungent odor, with young, sparsely pubescent branches; usually 1-2m tall, can sometimes reach 6m. The leaves are imparipinnate, formed by 13-17 (sometimes 7) elliptical segments, pubescent or subglabrous, carried by a 2 mm petiole; the stipules are linear, 3-4 mm long and precociously caducous. The numerous papilionaceous flowers, purplish-purple in color, are collected in linear spiciform racemes. The legumes, dotted-glandular and 7-9 mm long, contain 1 or 2 seeds. It can be confused with young plants or new shoots of Robinia pseudoacacia, which, however, are odorless and with hairless and thorny branches.
It is common broadleaved tree species in Europe, found from Scandinavia to the Iberian Peninsula. Oak trees can live for more than 1000 years and grow to be 30 to 40m in height. The wood from oaks is hard and durable and has been valued for centuries. Recently, concerns have arisen about the fate of oaks in the face of Acute Oak Decline, a little understood syndrome. This species, as well as other oaks, is very variable morphologically, and can naturally hybridise, generating individuals showing intermediate traits or the prevalence of one, so that it can be difficult to characterise them unequivocally by observations alone. The barks are grey, fissured, forming rectangular elongate blocks and thicker. The leaves are simple, obovate-oblong and deeply and irregularly lobed, with a short stalk (2-7mm). This oak is monoecious and wind-pollinated, with drooping male flowers in yellow catkins about 5cm long and inconspicuous globular female flowers of 1mm at terminal shots, which appear just after the first leaves have flushed. The fruits are the acorns, which are often in pairs and sit in scaly cups on the ends of long stalks. The acorns are very variable in size and shape rounded with olive-green longitudinal stripes visible when fresh. Mammals and birds are important for the seed dispersal, especially the Eurasian jay (Garrulus glandarius) which can be considered the primary propagator. Q. robur occur at many sites as a main component of temperate deciduous mixed forests, and they share several common characteristics. It is vigorous trees with a large ecological amplitude, although they prefer fertile and moist soils, and are able to dominate forests in number and size at low-mid elevations. As this tree do not come into leaf until relatively late in the year (late April to early May), late frost damage is rarely a problem, unless the temperatures reach -3°C killing new foliage. The tendency is for Q. robur to grow on heavier soils in more continental climates, in wet lowlands and damp areas by streams and rivers, tolerating periodic flooding. It is light-demanding and its canopies permit a good deal of light to pass through to the undergrowth, promoting the regeneration of many tree species and enriching forest diversity. These oaks rarely form pure forests under natural conditions. Their substantial competitor is represented by beech (Fagus sylvatica) and in a minor way other shade or half-shade trees, in the presence of which the oaks are unable to predominate as they are at a disadvantage. Typically, oaks dominate in damp to wet and nutrient-rich soils, where they occur principally with hornbeam (Carpinus betulus) and other deciduous tree species such as ash (Fraxinus excelsior, Fraxinus angustifolia), maple (Acer campestre, Acer platanoides) and small-leaved lime (Tilia cordata). In these oak-hornbeam forest communities, assigned to the Carpinion betuli alliance, beech is out of its range, or replaced as the soils are relatively dry and warm or too wet.